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Studies in Medicine, Homburg/Saar and Heidelberg |
| 1964 – 1966: Research Assistant, Anatomic Medical Faculty, Heidelberg University | |
| 1966 – 1972: Postdoc, Toronto University and Rockefeller University, New York | |
| 1972 – 1975 Professor of Anatomy, Heidelberg University | |
| 1976 – 2003: Head of Department, Department of Cytobiology and Cytopathology, Philipps University Marburg | |
| 1993 – 1995 and 1999 – 2000: Dean of the Medical Faculty, Philipps University Marburg | |
| 2000 – 2003: President of Philipps University Marburg | |
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