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1960 - 1967 Studies in Biologie, Chemistry, Physics und Philosophy at the University Frankfurt/M |
| 1970 Assistent professor at the Department of Zoology, University Zürich, Swiss | |
| 1973 - 1974 Research Fellow Department of Biology, Yale University, USA | |
| 1974 - 2005 Professor (chair) and 1986 - 2005 Director Department of Zoology, University Zürich, Swiss, since 2005: Emeritus | |
| 1988 - 1994 Andrew Dickson White Professor (at Large), Cornell University, USA | |
| 1990 - 2008 Non-resident Permanent Fellow Institute for Advanced Study, Berlin | |
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